The name is absent



KINETICS ON THE MICROBIAL SCALE

51


ASSUME (I0) , PYRUVATE , IS A FIXED INPUT PARAMETER

EQUATIONS

UNKNOWN

intermediates , ETC.

Г ≡ 5Γ                                 I

Ii

9

r = f,(I0)(On)- b1 (I0)(Rn)

r

I

r * f2(I, )(I9)- ⅛(12)

r≡ VI0W- ⅛(I9)(Rn)

9

IO

IO

FROM ELECTRON TRANSP.

12

13

GRAND TOTAL

22 EQ1NS

23 UNKNOWNS

fig.7-Determinacy in the krebs cycle and in
IN THE ENTIRE REACTION SYSTEM

Presumably there is an enzyme factory which can synthesize I8 at
arbitrarily small rate r√ I8 is also lost by diffusion from the system at
rate MI8, where M is a small mass transfer coefficient. The steady state
value for l8, rs∕M, is the ratio of two small numbers, and can presumably
be whatever the cell desires. The important fact is that the overall rate

FIG. 8- MODIFIED KREBS


CYCLE (SIMPLIFIED)

r is directly proportional to rs∕M in the steady state.

Since normally M would be fixed by cell morphology, the overall rate
of ATP formation can be controlled by the cell merely by changing the
small rate rs. This is the principle of the power amplifier and is charac-
teristic of cyclic reaction paths. It will be the subject of a demonstration
in the next paper.



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